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1. A small distinguishing mark on [the letters] "i" or "j". It was originally a larger mark, first appearing in Latin manuscripts, but was shortened when Roman-style typefaces were introduced. It is thought to be derived from Iota, [the smallest] letter in the Greek alphabet, which was used as a diacritic below other vowels in [ancient Greek] texts; others say it is derived from yodh, the smallest letter in Hebrew and Aramic alphabets. 2. A very small part or quantity; a particle, jot, or whit.
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