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One's [favorite person]; the one you love most. In Old English, the pupil of the eye (the round, dark center) was called the 'apple'. It was thought that the pupil was a round object much like an apple (a [piece of] fruit). When you look at someone, their [reflection] appears in your pupil. So if someone is the 'apple of your eye', he or she is someone that you look at a lot and enjoy seeing.
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