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1) To be called out for random and often awkward comments, with often hilarious results. 2) to be out of place or uncalled for. 3) to overreact in a manner that provokes uncalled for or awkward reactions 4) It is a phrase that came to be popular mainly in the northern areas of USA and more prominently in certain Canadian cities (Toronto, Ottawa & Montreal). Popular belief is that it came from the term Man-trany, which it self is a very uncalled and awkward term to use. The word "fill" was added in later as to fill in a blank space in a hypothetical sentence where " Man-trany" would be uncalled for.
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