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1. In machines, the amount of time, when operating a bank of machines, between any one of them failing. Not lifetime, as hard drive manufacturers want you to think. It is the average amount of time between failures of indivdual parts when talking about a system. 2. In biology, the average lifespan of each individual organism which has a particular set of genes or "environment of luck". An "environment of luck" can be something like all people who fly on a plane, living around individuals who want you dead in the worst way,all people who smoke, or some other demographic.
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